/* Honey Odyssey. Visual system: a modern natural-history museum.
   Light "gallery walls" (pale sage paper), dark specimen vitrines, one accent: pollen.
   Shape rule: media, vitrines and panels are square (radius 0); interactive controls are full pills. */

@font-face{font-family:"Newsreader";font-style:normal;font-display:swap;font-weight:200 800;src:url(fonts/newsreader-latin-opsz-normal.woff2) format("woff2-variations");unicode-range:U+0000-00FF,U+0131,U+0152-0153,U+02BB-02BC,U+02C6,U+02DA,U+02DC,U+0304,U+0308,U+0329,U+2000-206F,U+20AC,U+2122,U+2191,U+2193,U+2212,U+2215,U+FEFF,U+FFFD}
@font-face{font-family:"Newsreader";font-style:normal;font-display:swap;font-weight:200 800;src:url(fonts/newsreader-latin-ext-opsz-normal.woff2) format("woff2-variations");unicode-range:U+0100-02BA,U+02BD-02C5,U+02C7-02CC,U+02CE-02D7,U+02DD-02FF,U+1E00-1E9F,U+1EF2-1EFF,U+20A0-20AB,U+20AD-20C0,U+2C60-2C7F,U+A720-A7FF}
@font-face{font-family:"Newsreader";font-style:italic;font-display:swap;font-weight:200 800;src:url(fonts/newsreader-latin-opsz-italic.woff2) format("woff2-variations");unicode-range:U+0000-00FF,U+0131,U+0152-0153,U+02BB-02BC,U+02C6,U+02DA,U+02DC,U+0304,U+0308,U+0329,U+2000-206F,U+20AC,U+2122,U+2191,U+2193,U+2212,U+2215,U+FEFF,U+FFFD}
@font-face{font-family:"Newsreader";font-style:italic;font-display:swap;font-weight:200 800;src:url(fonts/newsreader-latin-ext-opsz-italic.woff2) format("woff2-variations");unicode-range:U+0100-02BA,U+02BD-02C5,U+02C7-02CC,U+02CE-02D7,U+02DD-02FF,U+1E00-1E9F,U+1EF2-1EFF,U+20A0-20AB,U+20AD-20C0,U+2C60-2C7F,U+A720-A7FF}
@font-face{font-family:"Hanken Grotesk";font-style:normal;font-display:swap;font-weight:100 900;src:url(fonts/hanken-grotesk-latin-wght-normal.woff2) format("woff2-variations");unicode-range:U+0000-00FF,U+0131,U+0152-0153,U+02BB-02BC,U+02C6,U+02DA,U+02DC,U+0304,U+0308,U+0329,U+2000-206F,U+20AC,U+2122,U+2191,U+2193,U+2212,U+2215,U+FEFF,U+FFFD}
@font-face{font-family:"Hanken Grotesk";font-style:normal;font-display:swap;font-weight:100 900;src:url(fonts/hanken-grotesk-latin-ext-wght-normal.woff2) format("woff2-variations");unicode-range:U+0100-02BA,U+02BD-02C5,U+02C7-02CC,U+02CE-02D7,U+02DD-02FF,U+1E00-1E9F,U+1EF2-1EFF,U+20A0-20AB,U+20AD-20C0,U+2C60-2C7F,U+A720-A7FF}

@font-face{font-family:"Newsreader";font-style:normal;font-display:swap;font-weight:200 800;src:url(fonts/literata-cyrillic-opsz-normal.woff2) format("woff2-variations");unicode-range:U+0301,U+0400-045F,U+0490-0491,U+04B0-04B1,U+2116}
@font-face{font-family:"Newsreader";font-style:italic;font-display:swap;font-weight:200 800;src:url(fonts/literata-cyrillic-opsz-italic.woff2) format("woff2-variations");unicode-range:U+0301,U+0400-045F,U+0490-0491,U+04B0-04B1,U+2116}

:root{
  --paper:#eef0e7; --paper-2:#e3e7da; --paper-3:#d6dccb;
  --ink:#15201a; --ink-2:#38463d; --ink-3:#56635a;
  --line:rgba(21,32,26,.16); --line-2:rgba(21,32,26,.32);
  --pollen:#f2b705; --pollen-2:#ffd34d; --pollen-ink:#7a5300; --on-pollen:#1d1503;
  --vitrine:#0a0c0b; --vitrine-ink:#e8eadf; --vitrine-ink-2:#a3aa9c;
  --nodata:#d3d8c8;
  --c1:#f7ebc1; --c2:#f2d26f; --c3:#e8b11c; --c4:#c78a0a; --c5:#93600a; --c6:#583808;
  --serif:"Newsreader",ui-serif,Georgia,"Hiragino Mincho ProN","Yu Mincho",serif;
  --sans:"Hanken Grotesk",ui-sans-serif,system-ui,-apple-system,"Segoe UI","Hiragino Sans","Noto Sans JP",sans-serif;
  --ease:cubic-bezier(.16,1,.3,1);
  --gut:clamp(16px,4vw,48px);
  --max:1360px;
  color-scheme:light dark;
}
@media (prefers-color-scheme:dark){:root:not([data-theme="light"]){
  --paper:#0e1511; --paper-2:#141d18; --paper-3:#1c2721;
  --ink:#e7eadf; --ink-2:#bec5b6; --ink-3:#97a093;
  --line:rgba(231,234,223,.14); --line-2:rgba(231,234,223,.3);
  --pollen-ink:#f4c94a; --vitrine:#050706; --nodata:#26312a;
  --c1:#3b2f12; --c2:#6b4f10; --c3:#a8780c; --c4:#dba21a; --c5:#f2c443; --c6:#ffe49a;
}}
:root[data-theme="dark"]{
  --paper:#0e1511; --paper-2:#141d18; --paper-3:#1c2721;
  --ink:#e7eadf; --ink-2:#bec5b6; --ink-3:#97a093;
  --line:rgba(231,234,223,.14); --line-2:rgba(231,234,223,.3);
  --pollen-ink:#f4c94a; --vitrine:#050706; --nodata:#26312a;
  --c1:#3b2f12; --c2:#6b4f10; --c3:#a8780c; --c4:#dba21a; --c5:#f2c443; --c6:#ffe49a;
}

*,*::before,*::after{box-sizing:border-box}
html{-webkit-text-size-adjust:100%;scroll-behavior:smooth;scroll-padding-top:80px}
@media (prefers-reduced-motion:reduce){html{scroll-behavior:auto}}
body{margin:0;background:var(--paper);color:var(--ink);font:400 17px/1.6 var(--sans);font-feature-settings:"ss01";text-rendering:optimizeLegibility;-webkit-font-smoothing:antialiased;overflow-x:hidden}
img,svg{display:block;max-width:100%}
a{color:inherit;text-decoration-thickness:1px;text-underline-offset:.2em}
a:hover{text-decoration-color:var(--pollen)}
:focus-visible{outline:2px solid var(--pollen);outline-offset:3px}
.sr{position:absolute!important;width:1px;height:1px;overflow:hidden;clip:rect(0 0 0 0);white-space:nowrap}
.skip{position:absolute;inset-inline-start:12px;top:-60px;z-index:100;background:var(--ink);color:var(--paper);padding:10px 16px;border-radius:999px}
.skip:focus{top:12px}
.wrap{max-width:var(--max);margin:0 auto;padding:0 var(--gut)}
.serif{font-family:var(--serif)}
i,em,.latin{font-family:var(--serif);font-style:italic}
.num{font-variant-numeric:tabular-nums lining-nums}

/* z-index scale: header 40, menu 50, tooltip 60, skip 100 */

/* ---------- Header ---------- */
.hdr{position:sticky;top:0;z-index:40;background:color-mix(in srgb,var(--paper) 86%,transparent);backdrop-filter:blur(14px) saturate(140%);-webkit-backdrop-filter:blur(14px) saturate(140%);border-bottom:1px solid transparent;transition:border-color .3s}
.hdr.is-scrolled{border-bottom-color:var(--line)}
.hdr .wrap{display:flex;align-items:center;gap:28px;height:68px}
.brand{display:flex;align-items:center;gap:10px;text-decoration:none;font-family:var(--serif);font-size:22px;font-weight:500;letter-spacing:-.01em;white-space:nowrap}
.brand svg{width:26px;height:26px}
.nav{display:flex;gap:4px;margin-inline-start:auto}
.nav a{font-size:15px;text-decoration:none;color:var(--ink-2);padding:8px 12px;border-radius:999px;transition:background .2s,color .2s;white-space:nowrap}
.nav a:hover{background:var(--paper-2);color:var(--ink)}
.lang{position:relative}
.lang>button,.menu-btn{font:500 14px var(--sans);color:var(--ink);background:transparent;border:1px solid var(--line-2);border-radius:999px;padding:7px 14px;cursor:pointer;display:flex;align-items:center;gap:6px}
.lang>button:hover,.menu-btn:hover{border-color:var(--ink)}
.lang-pop{position:absolute;inset-inline-end:0;top:calc(100% + 10px);background:var(--paper);border:1px solid var(--line-2);padding:8px;min-width:230px;box-shadow:0 18px 50px -18px rgba(21,32,26,.35);display:none;z-index:50}
.lang.is-open .lang-pop{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:2px}
.lang-pop a,.lang-pop span{display:flex;justify-content:space-between;gap:12px;padding:8px 10px;font-size:15px;text-decoration:none}
.lang-pop span{color:var(--ink-3)}
.lang-pop small{font-size:12px;color:var(--ink-3)}
.lang-pop a[aria-current]{background:var(--paper-2)}
.menu-btn{display:none}
@media (max-width:1060px){
  .nav{display:none}
  .menu-btn{display:flex;margin-inline-start:auto}
  .lang{margin-inline-start:0}
  .nav.is-open{display:flex;flex-direction:column;position:fixed;inset:68px 0 auto 0;background:var(--paper);padding:12px var(--gut) 24px;border-bottom:1px solid var(--line);z-index:50}
  .nav.is-open a{font-size:20px;padding:12px 4px;border-radius:0;border-bottom:1px solid var(--line)}
}
@media (min-width:1061px){.lang{margin-inline-start:8px}}

/* ---------- Buttons ---------- */
.btn{display:inline-flex;align-items:center;gap:10px;font:600 16px/1 var(--sans);text-decoration:none;padding:15px 24px;border-radius:999px;border:1px solid transparent;cursor:pointer;white-space:nowrap;transition:transform .25s var(--ease),background .25s,border-color .25s}
.btn:active{transform:translateY(1px) scale(.985)}
.btn-primary{background:var(--pollen);color:var(--on-pollen)}
.btn-primary:hover{background:var(--pollen-2)}
.btn-ghost{color:var(--ink);border-color:var(--line-2)}
.btn-ghost:hover{border-color:var(--ink)}
.btn svg{width:18px;height:18px;transition:transform .3s var(--ease)}
.btn:hover svg{transform:translateX(3px)}

/* ---------- Hero ---------- */
.hero{padding:clamp(28px,5vw,72px) 0 clamp(48px,7vw,104px)}
.hero .wrap{display:grid;grid-template-columns:minmax(0,11fr) minmax(0,13fr);gap:clamp(28px,5vw,80px);align-items:end}
.hero h1{font-family:var(--serif);font-weight:330;font-size:clamp(46px,6.1vw,92px);line-height:1.02;letter-spacing:-.035em;margin:0 0 28px;font-variation-settings:"opsz" 72}
.hero h1 em{font-weight:300;display:inline-block;padding-bottom:.08em}
.hero .lede{font-size:clamp(18px,1.6vw,21px);line-height:1.5;color:var(--ink-2);max-width:30ch;margin:0 0 34px}
.cta-row{display:flex;flex-wrap:wrap;gap:12px}
.hero-copy{padding-bottom:6px}
.hero-copy>*{animation:rise 1s var(--ease) both}
.hero-copy>*:nth-child(2){animation-delay:.08s}.hero-copy>*:nth-child(3){animation-delay:.16s}
@keyframes rise{from{opacity:0;transform:translateY(18px)}to{opacity:1;transform:none}}

.vitrine{background:var(--vitrine);color:var(--vitrine-ink);position:relative;overflow:hidden;margin:0}
.vitrine img{width:100%;height:100%;object-fit:cover}
.specimen-label{display:grid;grid-template-columns:auto 1fr;gap:4px 18px;font-size:13px;line-height:1.45;color:var(--ink-3);padding:14px 0 0;border-top:1px solid var(--line);margin-top:14px}
.specimen-label b{font-weight:600;color:var(--ink-2)}
.hero-figure .vitrine{aspect-ratio:3/2;animation:fade 1.4s var(--ease) both .1s}
@keyframes fade{from{opacity:0}to{opacity:1}}
.hero-figure .vitrine img{transform-origin:62% 38%}
@supports (animation-timeline:view()){
  @media (prefers-reduced-motion:no-preference){
    .hero-figure .vitrine img{animation:zoomin linear both;animation-timeline:view();animation-range:cover 50% cover 100%}
    @keyframes zoomin{from{transform:scale(1)}to{transform:scale(1.55)}}
  }
}
.vitrine-glare{position:absolute;inset:0;pointer-events:none;background:linear-gradient(115deg,rgba(255,255,255,.07),transparent 28%,transparent 72%,rgba(255,255,255,.04));box-shadow:inset 0 0 0 1px rgba(255,255,255,.06)}

/* ---------- Section rhythm ---------- */
section{position:relative}
.sec{padding:clamp(72px,10vw,150px) 0}
.sec-head{max-width:760px;margin-bottom:clamp(36px,5vw,64px)}
.h2{font-family:var(--serif);font-weight:350;font-size:clamp(36px,5vw,68px);line-height:1.04;letter-spacing:-.03em;margin:0 0 18px;font-variation-settings:"opsz" 60}
.h2 em{font-weight:320}
.sec-head p{font-size:19px;color:var(--ink-2);margin:0;max-width:58ch}
.sec-head .eyebrow,.eyebrow{font:600 12px/1 var(--sans);letter-spacing:.16em;text-transform:uppercase;color:var(--pollen-ink);margin:0 0 22px}
.reveal{opacity:0;transform:translateY(24px);transition:opacity .9s var(--ease),transform .9s var(--ease)}
.reveal.is-in{opacity:1;transform:none}
@media (prefers-reduced-motion:reduce){.reveal{opacity:1;transform:none;transition:none}}

/* ---------- Evidence chips ---------- */
.ev{display:inline-flex;align-items:center;gap:8px;font:500 13px/1.3 var(--sans);color:var(--ink-2);text-decoration:none}
.ev::before{content:"";width:10px;height:10px;flex:none;border:1.5px solid currentColor}
.ev-fact::before{background:var(--pollen);border-color:var(--pollen-ink);border-radius:50%}
.ev-estimate::before{border-radius:50%;background:linear-gradient(90deg,var(--pollen-ink) 50%,transparent 50%);border-color:var(--pollen-ink)}
.ev-observation::before{border-radius:50%}
.ev-reported::before{border-radius:0}
.ev-question::before{border-radius:50%;border-style:dashed}
.ev a{color:var(--ink-3)}
.src{font-size:13px;color:var(--ink-3);line-height:1.5}
.src a{color:var(--ink-3)}

/* ---------- Journey ---------- */
.journey{background:var(--paper-2)}
.journey-grid{display:grid;grid-template-columns:minmax(0,5fr) minmax(0,6fr);gap:clamp(24px,5vw,90px)}
.steps{counter-reset:step}
.step{min-height:78vh;display:flex;flex-direction:column;justify-content:center;padding:6vh 0;opacity:.32;transition:opacity .6s var(--ease)}
.step.is-active{opacity:1}
.step:first-child{min-height:60vh}
.step h3{font-family:var(--serif);font-weight:400;font-size:clamp(30px,3.2vw,44px);line-height:1.08;letter-spacing:-.02em;margin:0 0 16px}
.step p{font-size:19px;line-height:1.6;color:var(--ink-2);margin:0 0 18px;max-width:46ch}
.step .meta{display:flex;flex-direction:column;gap:6px}
.step-verb{font:600 13px/1 var(--sans);letter-spacing:.04em;color:var(--pollen-ink);margin-bottom:14px}
.stage{position:sticky;top:96px;height:calc(100dvh - 140px);min-height:460px}
.stage .vitrine{position:absolute;inset:0}
.layer{position:absolute;inset:0;opacity:0;transition:opacity .9s var(--ease),transform 1.4s var(--ease);transform:scale(1.04)}
.layer.is-on{opacity:1;transform:none}
.layer img{width:100%;height:100%;object-fit:cover}
.gauge{position:absolute;inset-inline-end:20px;bottom:20px;top:20px;width:76px;display:flex;flex-direction:column;align-items:center;gap:10px;z-index:2}
.gauge-tube{position:relative;flex:1;width:22px;border:1px solid rgba(232,234,223,.5);border-radius:999px;overflow:hidden;background:rgba(10,12,11,.55);backdrop-filter:blur(6px)}
.gauge-fill{position:absolute;inset-inline-start:0;inset-inline-end:0;bottom:0;height:var(--water,92%);background:linear-gradient(180deg,rgba(175,214,226,.95),rgba(120,170,190,.9));transition:height 1.2s var(--ease)}
.gauge-line{position:absolute;inset-inline-start:-6px;inset-inline-end:-6px;bottom:20%;border-top:1.5px solid var(--pollen)}
.gauge-cap{font:600 11px/1.25 var(--sans);text-align:center;color:var(--vitrine-ink);text-shadow:0 1px 8px rgba(0,0,0,.7)}
.gauge-cap span{display:block;color:var(--pollen-2);font-size:13px}
.stage-cap{position:absolute;inset-inline-start:0;inset-inline-end:110px;bottom:0;padding:18px 20px;font-size:13px;color:var(--vitrine-ink-2);background:linear-gradient(0deg,rgba(5,7,6,.85),transparent);z-index:2}
.codex{font-family:var(--serif);font-size:clamp(22px,2.1vw,28px);line-height:1.35;font-weight:350;margin:0 0 18px;padding-inline-start:20px;border-inline-start:2px solid var(--pollen);max-width:40ch;color:var(--ink)}
.journey-mobile-img{display:none}
@media (max-width:880px){
  .journey-grid{grid-template-columns:1fr}
  .stage{display:none}
  .step,.step:first-child{min-height:auto;opacity:1;padding:28px 0;border-top:1px solid var(--line)}
  .journey-mobile-img{display:block;aspect-ratio:4/3;margin:0 0 20px}
}

/* ---------- Hive ---------- */
.hive-grid{display:grid;grid-template-columns:minmax(0,8fr) minmax(0,4fr);gap:clamp(24px,4vw,56px);align-items:start}
.comb-wrap{background:var(--vitrine);padding:clamp(10px,2vw,22px)}
.comb{width:100%;height:auto}
.comb .cell{transition:opacity .45s var(--ease)}
.comb[data-active] .cell{opacity:.16}
.comb[data-active="eggs"] .z-eggs,.comb[data-active="larvae"] .z-larvae,.comb[data-active="capped"] .z-capped,.comb[data-active="pollen"] .z-pollen,.comb[data-active="nectar"] .z-nectar,.comb[data-active="honey"] .z-honey,.comb[data-active="drone"] .z-drone,.comb[data-active="queen"] .z-queen{opacity:1}
.comb[data-active] .z-queen{opacity:.16}
.comb[data-active="queen"] .z-queen{opacity:1}
.legend{display:flex;flex-wrap:wrap;gap:8px;margin:18px 0 0}
.legend button{font:500 14px/1 var(--sans);display:inline-flex;align-items:center;gap:8px;padding:9px 14px 9px 10px;border-radius:999px;border:1px solid var(--line-2);background:transparent;color:var(--ink);cursor:pointer;transition:background .2s,border-color .2s}
.legend button:hover{border-color:var(--ink)}
.legend button[aria-pressed="true"]{background:var(--ink);color:var(--paper);border-color:var(--ink)}
.legend i{width:14px;height:14px;display:inline-block;clip-path:polygon(25% 3%,75% 3%,100% 50%,75% 97%,25% 97%,0 50%)}
.hive-panel{position:sticky;top:96px}
.hive-panel h3{font-family:var(--serif);font-weight:400;font-size:34px;line-height:1.1;margin:0 0 12px;letter-spacing:-.02em}
.hive-panel p{color:var(--ink-2);margin:0 0 16px;font-size:18px}
.hive-photo{margin:28px 0 0}
.hive-photo .vitrine{aspect-ratio:16/9}
.hive-photo figcaption{font-size:13px;color:var(--ink-3);margin-top:10px}
.castes{display:grid;grid-template-columns:repeat(12,minmax(0,1fr));gap:clamp(12px,2vw,24px);margin-top:clamp(56px,7vw,96px)}
.caste{grid-column:span 4}
.caste:nth-child(2){margin-top:48px}
.caste:nth-child(3){margin-top:96px}
.caste .vitrine{aspect-ratio:4/3}
.caste h4{font-family:var(--serif);font-weight:400;font-size:28px;margin:16px 0 4px;letter-spacing:-.01em}
.caste .days{font:500 15px var(--sans);color:var(--ink-2)}
.caste .days b{font-family:var(--serif);font-weight:400;font-size:44px;color:var(--ink);letter-spacing:-.02em;margin-inline-end:6px}
.caste p{font-size:15px;color:var(--ink-3);margin:6px 0 0}
.castes-src{grid-column:1/-1;margin-top:8px}
@media (max-width:900px){
  .hive-grid{grid-template-columns:1fr}
  .hive-panel{position:static}
  .caste{grid-column:1/-1;margin-top:0!important}
}

/* ---------- Diversity ---------- */
.div-top{display:grid;grid-template-columns:minmax(0,7fr) minmax(0,4fr);gap:clamp(24px,5vw,72px);align-items:end;margin-bottom:clamp(36px,5vw,56px)}
.div-top figure{margin:0}
.div-top .vitrine{aspect-ratio:3/4;max-height:520px}
.div-top figcaption{font-size:13px;color:var(--ink-3);margin-top:10px}
.big-stat{font-family:var(--serif);font-weight:300;font-size:clamp(72px,11vw,168px);line-height:.9;letter-spacing:-.05em;margin:0 0 12px}
.big-stat small{font-size:.3em;letter-spacing:-.01em;font-weight:400;margin-inline-start:.12em}
.rail{display:grid;grid-auto-flow:column;grid-auto-columns:minmax(260px,22%);gap:18px;overflow-x:auto;scroll-snap-type:x mandatory;padding:4px var(--gut) 24px;margin:0 calc(-1 * var(--gut));scrollbar-width:thin;scrollbar-color:var(--line-2) transparent}
.rail:focus-visible{outline-offset:-2px}
.spec{scroll-snap-align:start;margin:0}
.spec .vitrine{aspect-ratio:1/1}
.spec .vitrine img{transition:transform 1.1s var(--ease)}
.spec:hover .vitrine img{transform:scale(1.07)}
.spec figcaption{padding-top:14px}
.spec .latin{font-size:22px;line-height:1.15;display:block}
.spec .common{font-size:14px;color:var(--ink-3);display:block;margin:2px 0 10px}
.spec p{font-size:15px;line-height:1.5;color:var(--ink-2);margin:0 0 8px}
.spec .nest{font:600 12px/1 var(--sans);letter-spacing:.06em;text-transform:uppercase;color:var(--pollen-ink)}
.rail-nav{display:flex;gap:8px;justify-content:flex-end;margin-top:-4px}
.rail-nav button{width:44px;height:44px;border-radius:999px;border:1px solid var(--line-2);background:transparent;color:var(--ink);cursor:pointer;display:grid;place-items:center}
.rail-nav button:hover{border-color:var(--ink)}
@media (max-width:900px){.div-top{grid-template-columns:1fr}.rail{grid-auto-columns:78%}.div-top .vitrine{aspect-ratio:4/3}}

/* ---------- Numbers ---------- */
.numbers{background:var(--paper-2)}
.num-grid{display:grid;grid-template-columns:minmax(0,7fr) minmax(0,5fr);gap:clamp(28px,5vw,80px)}
.figure-xl{font-family:var(--serif);font-weight:300;font-size:clamp(64px,9vw,136px);line-height:.92;letter-spacing:-.045em;margin:0}
.figure-xl small{font-size:.32em;letter-spacing:-.01em;font-weight:400}
.figure-label{font-size:19px;color:var(--ink-2);margin:12px 0 20px;max-width:40ch}
.meta-list{display:grid;grid-template-columns:auto 1fr;gap:4px 18px;font-size:14px;color:var(--ink-3);margin:0}
.meta-list dt{color:var(--ink-2);font-weight:600}
.meta-list dd{margin:0}
.chart{margin-top:36px}
.chart svg{width:100%;height:auto;overflow:visible}
.chart .axis text{font:500 12px var(--sans);fill:var(--ink-3)}
.chart .axis line{stroke:var(--line)}
.chart .area{fill:var(--pollen);opacity:.18}
.chart .ln{fill:none;stroke:var(--pollen-ink);stroke-width:2.2;stroke-linejoin:round}
.chart .dot{fill:var(--pollen-ink)}
.chart.draw .ln{stroke-dasharray:var(--len);stroke-dashoffset:var(--len);transition:stroke-dashoffset 2.2s var(--ease)}
.chart.draw.is-in .ln{stroke-dashoffset:0}
@media (prefers-reduced-motion:reduce){.chart.draw .ln{stroke-dasharray:none;stroke-dashoffset:0;transition:none}}
.chart-note{font-size:13px;color:var(--ink-3);margin-top:8px}
.side-stats{display:flex;flex-direction:column;gap:clamp(28px,4vw,44px)}
.side-stat{padding-top:20px;border-top:1px solid var(--line-2)}
.side-stat .v{font-family:var(--serif);font-weight:350;font-size:clamp(44px,5vw,64px);line-height:1;letter-spacing:-.03em}
.side-stat .v small{font-size:.4em;font-weight:400}
.side-stat .l{font-size:16px;color:var(--ink-2);margin:8px 0 10px}
.honest{background:var(--paper);padding:22px 22px 20px;border-inline-start:3px solid var(--pollen)}
.honest h3{font-family:var(--serif);font-weight:400;font-size:24px;margin:0 0 8px;letter-spacing:-.01em}
.honest p{margin:0;font-size:15.5px;color:var(--ink-2)}
@media (max-width:900px){.num-grid{grid-template-columns:1fr}}

/* ---------- Map ---------- */
.map-shell{display:grid;grid-template-columns:minmax(0,9fr) minmax(0,3fr);gap:clamp(20px,3vw,40px);align-items:start}
.map-tools{display:flex;flex-wrap:wrap;gap:8px;margin-bottom:16px}
.chip{font:500 14px/1 var(--sans);padding:10px 16px;border-radius:999px;border:1px solid var(--line-2);background:transparent;color:var(--ink);cursor:pointer}
.chip[aria-pressed="true"]{background:var(--ink);color:var(--paper);border-color:var(--ink)}
.chip:disabled{color:var(--ink-3);border-style:dashed;cursor:not-allowed}
.chip:disabled small{font-size:12px;margin-inline-start:6px}
.map-box{position:relative;aspect-ratio:2/1.04;background:var(--paper-2)}
.map-box svg{width:100%;height:100%}
.map-box .ctry{stroke:var(--paper);stroke-width:.5;transition:fill .5s var(--ease)}
.map-box .ctry:hover,.map-box .ctry:focus{stroke:var(--ink);stroke-width:1.2;outline:none}
.map-box .sphere{fill:none;stroke:var(--line)}
.map-box .grat{fill:none;stroke:var(--line);stroke-width:.4}
.map-loading{position:absolute;inset:0;display:grid;place-items:center;color:var(--ink-3);font-size:14px}
.map-loading::before{content:"";position:absolute;inset:0;background:linear-gradient(100deg,transparent 30%,color-mix(in srgb,var(--paper) 60%,transparent) 50%,transparent 70%);background-size:200% 100%;animation:shimmer 1.6s linear infinite}
@keyframes shimmer{from{background-position:120% 0}to{background-position:-80% 0}}
.tip{position:absolute;z-index:60;pointer-events:none;background:var(--ink);color:var(--paper);padding:10px 12px;font-size:13px;line-height:1.4;min-width:170px;transform:translate(-50%,calc(-100% - 12px));opacity:0;transition:opacity .15s}
.tip.is-on{opacity:1}
.tip b{display:block;font-size:14px;margin-bottom:2px}
.scale{display:flex;align-items:center;gap:0;margin-top:14px;flex-wrap:wrap}
.scale span{display:flex;flex-direction:column;font-size:12px;color:var(--ink-3);min-width:64px}
.scale span::before{content:"";height:10px;background:var(--sw);margin-bottom:6px}
.scale .nd{margin-inline-start:16px}
.scale .nd::before{background:repeating-linear-gradient(45deg,var(--nodata) 0 3px,transparent 3px 6px);border:1px solid var(--line)}
.rank h3{font-family:var(--serif);font-weight:400;font-size:24px;margin:0 0 4px}
.rank .src{margin:0 0 14px}
.rank ol{list-style:none;margin:0;padding:0;counter-reset:r}
.rank li{counter-increment:r;display:grid;grid-template-columns:26px 1fr auto;gap:2px 8px;align-items:baseline;padding:7px 0;font-size:15px}
.rank li::before{content:counter(r);color:var(--ink-3);font-size:13px}
.rank li .bar{grid-column:2/4;height:2px;margin-top:3px;background:var(--c3);width:var(--w)}
.rank li .v{font-variant-numeric:tabular-nums;color:var(--ink-2)}
.map-notes{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:18px 40px;margin-top:28px;font-size:14px;color:var(--ink-3)}
.map-notes p{margin:0}
@media (max-width:900px){.map-tools{flex-wrap:nowrap;overflow-x:auto;margin:0 calc(-1*var(--gut)) 16px;padding:0 var(--gut);scrollbar-width:none}.chip{flex:none}.map-shell{grid-template-columns:1fr}.map-notes{grid-template-columns:1fr}}

/* ---------- Journeys list ---------- */
.ways{border-top:1px solid var(--line-2)}
.way{display:grid;grid-template-columns:minmax(0,6fr) minmax(0,5fr) auto;gap:24px;align-items:center;padding:clamp(22px,3vw,34px) 0;border-bottom:1px solid var(--line-2);text-decoration:none;position:relative}
.way h3{font-family:var(--serif);font-weight:350;font-size:clamp(30px,4vw,54px);letter-spacing:-.03em;line-height:1.05;margin:0;transition:transform .5s var(--ease)}
a.way:hover h3{transform:translateX(10px)}
.way p{margin:0;color:var(--ink-2);font-size:16px}
.way .status{font:600 13px var(--sans);white-space:nowrap;color:var(--ink-3)}
a.way .status{color:var(--pollen-ink)}
@media (max-width:800px){.way{grid-template-columns:1fr;gap:8px}}

/* ---------- Method ---------- */
.method-grid{display:grid;grid-template-columns:minmax(0,5fr) minmax(0,7fr);gap:clamp(28px,5vw,90px)}
.evidence{list-style:none;margin:0;padding:0;display:grid;align-content:start;grid-template-columns:repeat(2,minmax(0,1fr));gap:0 32px}
.evidence li{padding:18px 0;border-top:1px solid var(--line-2)}
.evidence li .ev{font-size:16px;color:var(--ink);font-weight:600;margin-bottom:6px}
.evidence li p{margin:0;font-size:15px;color:var(--ink-2)}
.principles{margin:0;padding:0;list-style:none;counter-reset:p}
.principles li{font-size:17px;color:var(--ink-2);padding:0 0 16px 0;margin:0 0 16px;border-bottom:1px solid var(--line)}
.principles b{color:var(--ink);font-weight:600}
@media (max-width:800px){.method-grid,.evidence{grid-template-columns:1fr}}

/* ---------- References ---------- */
.refs{background:var(--paper-2);padding:clamp(56px,7vw,96px) 0}
.refs h2{font-family:var(--serif);font-weight:400;font-size:32px;margin:0 0 24px}
.refs ol{columns:2 420px;column-gap:56px;margin:0;padding-inline-start:22px;font-size:14.5px;color:var(--ink-2)}
.refs li{break-inside:avoid;margin:0 0 12px;padding-inline-start:4px}
.refs li:target{background:color-mix(in srgb,var(--pollen) 22%,transparent)}

/* ---------- Footer ---------- */
.ftr{padding:56px 0 40px;border-top:1px solid var(--line)}
.ftr .wrap{display:grid;grid-template-columns:minmax(0,5fr) repeat(2,minmax(0,3fr));gap:32px}
.ftr p{margin:0 0 10px;font-size:14px;color:var(--ink-3)}
.ftr h4{font:600 14px var(--sans);margin:0 0 12px}
.ftr ul{list-style:none;margin:0;padding:0;font-size:14px;color:var(--ink-2)}
.ftr li{margin:0 0 8px}
.ftr .motto{font-family:var(--serif);font-size:28px;color:var(--ink);line-height:1.2;margin-bottom:14px;letter-spacing:-.01em}
@media (max-width:800px){.ftr .wrap{grid-template-columns:1fr}}

/* ---------- Simple pages ---------- */
.page{padding:clamp(40px,6vw,88px) 0}
.page h1{font-family:var(--serif);font-weight:350;font-size:clamp(40px,5.5vw,72px);letter-spacing:-.03em;line-height:1.04;margin:0 0 20px}
.page .lede{font-size:19px;color:var(--ink-2);max-width:60ch;margin:0 0 40px}
.credit-list{display:grid;grid-template-columns:repeat(auto-fill,minmax(300px,1fr));gap:28px}
.credit{margin:0}
.credit .vitrine{aspect-ratio:3/2}
.credit figcaption{font-size:14px;color:var(--ink-2);padding-top:10px;line-height:1.5}
.credit figcaption b{display:block;color:var(--ink);font-weight:600}
.notfound{min-height:60vh;display:flex;flex-direction:column;justify-content:center}

@media (max-width:760px){
  body{font-size:16px}
  .hero .wrap{grid-template-columns:1fr}
  .hero-figure{order:-1}
  .hero h1{font-size:clamp(44px,13vw,64px)}
  .btn{padding:14px 20px;font-size:15px}
  .step p,.sec-head p{font-size:17px}
}

/* ---------- Catalogue and species pages ---------- */
.crumbs{display:flex;flex-wrap:wrap;gap:8px;font-size:14px;color:var(--ink-3);margin:0 0 28px}
.crumbs a{color:var(--ink-3)}
.page h1 em,.page h1 i{font-weight:320}
.spec a.latin{text-decoration:none}
.spec a.latin:hover{text-decoration:underline;text-decoration-color:var(--pollen)}
.coverage{display:grid;grid-template-columns:minmax(0,1fr);gap:10px;max-width:640px;margin:0 0 36px}
.cov-bar{height:6px;background:var(--paper-3);position:relative}
.cov-bar span{position:absolute;inset:0 auto 0 0;width:var(--p);background:var(--pollen-ink);min-width:4px}
.coverage p{margin:0;font-size:15px;color:var(--ink-2)}
.fam-filter{margin-bottom:8px}
.fam-group{padding:44px 0 8px;border-top:1px solid var(--line-2);margin-top:28px}
.fam-group[hidden]{display:none}
.fam-head{display:grid;grid-template-columns:minmax(0,4fr) minmax(0,7fr) auto;gap:24px;align-items:baseline;margin-bottom:28px}
.fam-head h2{font-family:var(--serif);font-weight:380;font-size:clamp(30px,3.4vw,44px);letter-spacing:-.02em;margin:0}
.fam-head p{margin:0;color:var(--ink-2);font-size:16px;max-width:60ch}
.fam-head .count{font-family:var(--serif);font-size:28px;color:var(--ink-3)}
@media (max-width:760px){.fam-head{grid-template-columns:1fr auto}.fam-head p{grid-column:1/-1;grid-row:2}}
.sp-grid{display:grid;grid-template-columns:repeat(auto-fill,minmax(220px,1fr));gap:28px 20px}
.sp-card{text-decoration:none;display:block}
.sp-card .vitrine{aspect-ratio:4/3}
.sp-card .vitrine img{transition:transform 1s var(--ease)}
.sp-card:hover .vitrine img{transform:scale(1.06)}
.sp-card .latin{display:block;font-size:21px;line-height:1.2;margin-top:12px}
.sp-card .common{display:block;font-size:14px;color:var(--ink-2);margin-top:2px}
.sp-card .fam{display:block;font:600 11px/1 var(--sans);letter-spacing:.1em;text-transform:uppercase;color:var(--ink-3);margin-top:8px}
.species .sp-hero{display:grid;grid-template-columns:minmax(0,5fr) minmax(0,7fr);gap:clamp(24px,5vw,72px);align-items:end;margin-bottom:clamp(40px,6vw,80px)}
.sp-id h1{font-family:var(--serif);font-weight:320;font-size:clamp(44px,5.6vw,84px);line-height:1.05;letter-spacing:-.03em;margin:0;padding-bottom:.06em}
.fam-line{font:600 12px/1 var(--sans);letter-spacing:.16em;text-transform:uppercase;color:var(--pollen-ink);margin:0 0 18px}
.authority{margin:6px 0 0;color:var(--ink-3);font-size:15px}
.common-line{font-size:20px;color:var(--ink);margin:18px 0 0}
.sp-id .lede{margin:18px 0 0;font-size:18.5px}
.sp-photo{margin:0}
.sp-photo .vitrine{aspect-ratio:3/2}
.sp-photo figcaption,.sp-second figcaption{font-size:13px;color:var(--ink-3);margin-top:10px}
.sp-body{display:grid;grid-template-columns:minmax(0,4fr) minmax(0,8fr);gap:clamp(24px,5vw,80px);align-items:start}
.sp-quick{position:sticky;top:96px}
.sp-quick dl{margin:0;display:grid;grid-template-columns:1fr;gap:0}
.sp-quick dt{font:600 12px/1 var(--sans);letter-spacing:.08em;text-transform:uppercase;color:var(--ink-3);padding-top:16px;border-top:1px solid var(--line);margin-top:14px}
.sp-quick dt:first-child{margin-top:0}
.sp-quick dd{margin:8px 0 0;font-size:17px;line-height:1.45}
.sp-quick dd small{display:block;font-size:13px;color:var(--ink-3);margin-top:2px}
.h3{font-family:var(--serif);font-weight:380;font-size:clamp(28px,3vw,40px);letter-spacing:-.02em;line-height:1.1;margin:0 0 24px}
.fact-block{padding:22px 0 6px;border-top:1px solid var(--line-2)}
.fact-block h3{font:600 13px/1 var(--sans);letter-spacing:.08em;text-transform:uppercase;color:var(--ink-2);margin:0 0 14px}
.fact-block ul{list-style:none;margin:0;padding:0}
.fact-block li{margin:0 0 18px}
.fact-block li p{margin:0 0 6px;font-size:18px;line-height:1.55;max-width:62ch}
.sp-second{margin:36px 0 0}
.sp-second .vitrine{aspect-ratio:16/10}
.sp-map{margin:clamp(56px,7vw,96px) 0 0;padding:clamp(36px,5vw,60px) 0;border-top:1px solid var(--line-2)}
.sp-map-grid{display:grid;grid-template-columns:minmax(0,4fr) minmax(0,8fr);gap:clamp(24px,4vw,64px);align-items:start}
.sp-map p{color:var(--ink-2);font-size:16px;margin:0 0 14px;max-width:48ch}
.sp-map .big{font-family:var(--serif);font-size:clamp(44px,5vw,64px);line-height:1;letter-spacing:-.03em;color:var(--ink);margin:20px 0 14px}
.sp-map .big small{font-size:.35em;font-family:var(--sans);letter-spacing:0;color:var(--ink-2)}
.toplist{list-style:none;margin:0 0 18px;padding:0;font-size:15px}
.toplist li{display:flex;justify-content:space-between;gap:12px;padding:6px 0;border-bottom:1px solid var(--line)}
.toplist .v{font-variant-numeric:tabular-nums;color:var(--ink-2)}
.caveat{font-size:14px!important;color:var(--ink-3)!important;border-inline-start:2px solid var(--pollen);padding-inline-start:12px}
.sp-map-box{position:relative;aspect-ratio:1024/404;background:var(--paper-2)}
.sp-map-box svg{width:100%;height:100%}
.sp-map-box .land{fill:var(--paper-3);stroke:var(--paper);stroke-width:.4}
.sp-map-box .sphere{fill:none;stroke:var(--line)}
.sens{display:grid;place-items:center;text-align:center;padding:24px;height:100%;color:var(--ink-2);font-size:15px}
.sp-rel{margin-top:clamp(48px,6vw,80px);padding-top:clamp(32px,4vw,48px);border-top:1px solid var(--line-2)}
.pn{display:grid;grid-template-columns:1fr 1fr;gap:20px;margin:clamp(40px,5vw,64px) 0 0}
.pn a{display:block;text-decoration:none;padding:20px 0;border-top:1px solid var(--ink)}
.pn a.next{text-align:end}
.pn small{display:block;font-size:13px;color:var(--ink-3);margin-bottom:6px}
.pn i{font-size:clamp(22px,2.4vw,30px)}
.sp-src{margin-top:clamp(48px,6vw,80px);padding:clamp(32px,4vw,48px) clamp(16px,3vw,40px);background:var(--paper-2)}
.sp-src ol{margin:0 0 20px;padding-inline-start:22px;font-size:14.5px;color:var(--ink-2);columns:2 380px;column-gap:48px}
.sp-src li{break-inside:avoid;margin-bottom:10px}
.sp-src li:target{background:color-mix(in srgb,var(--pollen) 22%,transparent)}
@media (max-width:900px){.species .sp-hero,.sp-body,.sp-map-grid{grid-template-columns:1fr}.sp-photo{order:-1}.sp-quick{position:static}}

/* ---------- Plants ---------- */
.cal-wrap{position:relative;overflow-x:auto;margin:8px 0 0;border-top:1px solid var(--line-2)}
.cal{border-collapse:collapse;width:100%;min-width:760px;font-size:14px}
.cal thead th{font:600 12px/1 var(--sans);letter-spacing:.06em;text-transform:uppercase;color:var(--ink-3);padding:14px 0 10px;text-align:center}
.cal thead th:first-child{text-align:start}
.cal tbody th{text-align:start;font-weight:400;padding:10px 16px 10px 0;width:30%;border-top:1px solid var(--line)}
.cal tbody th a{font-size:17px;text-decoration:none}
.cal tbody th a:hover{text-decoration:underline;text-decoration-color:var(--pollen)}
.cal tbody th small{display:block;color:var(--ink-3);font-size:13px}
.cal td{border-top:1px solid var(--line);padding:0 1px;height:44px}
.cal td.on{background:linear-gradient(var(--pollen),var(--pollen)) center/100% 14px no-repeat}
.cal tbody tr:hover td.on{background-image:linear-gradient(var(--pollen-ink),var(--pollen-ink))}
.inv{font:600 10px/1 var(--sans);letter-spacing:.08em;text-transform:uppercase;color:var(--pollen-ink);border:1px solid currentColor;padding:2px 5px;border-radius:999px;margin-inline-start:4px}
.mstrip{display:grid;grid-template-columns:repeat(12,1fr);gap:2px;max-width:280px;margin-top:2px}
.mstrip span{font:600 10px/1 var(--sans);text-align:center;padding:5px 0;background:var(--paper-2);color:var(--ink-3)}
.mstrip span.on{background:var(--pollen);color:var(--on-pollen)}
.links{list-style:none;margin:0;padding:0}
.links li{margin:0 0 16px}
.links li a{font-size:20px}
.links li p{margin:2px 0 6px;font-size:16px;color:var(--ink-2)}
.xlink{margin:18px 0 0}
.xlink a{display:inline-flex;gap:8px;align-items:center;font-weight:600;text-decoration:none;border-bottom:1px solid var(--pollen)}
.xlink svg{width:16px;height:16px}

/* ---------- Editorial pages ---------- */
.process{list-style:none;margin:0;padding:0;display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:0;counter-reset:pr;border-top:1px solid var(--ink)}
.process li{padding:22px 22px 8px 0;counter-increment:pr}
.process li::before{content:counter(pr);font-family:var(--serif);font-size:44px;line-height:1;color:var(--pollen-ink);display:block;margin-bottom:12px}
.process h3{font-family:var(--serif);font-weight:400;font-size:24px;margin:0 0 8px}
.process p{margin:0;color:var(--ink-2);font-size:16px}
@media (max-width:900px){.process{grid-template-columns:1fr 1fr}}
@media (max-width:560px){.process{grid-template-columns:1fr}}
.srclist{margin:0;padding-inline-start:24px;columns:2 420px;column-gap:48px;font-size:14.5px;color:var(--ink-2)}
.srclist li{break-inside:avoid;margin-bottom:10px}
.cform{display:grid;gap:22px;max-width:620px}
.fld{display:grid;gap:8px}
.fld label{font-weight:600;font-size:15px}
.fld input,.fld textarea{font:400 17px/1.5 var(--sans);color:var(--ink);background:var(--paper);border:1px solid var(--line-2);padding:12px 14px;border-radius:0}
.fld input:focus,.fld textarea:focus{outline:2px solid var(--pollen);outline-offset:0;border-color:var(--ink)}
.fld input::placeholder{color:var(--ink-3)}
.fld small{font-size:13px;color:var(--ink-3)}
.fld .err{color:#a1361a}
.hp{position:absolute;left:-9999px;width:1px;height:1px;overflow:hidden}
.cstatus{margin:0;font-size:15px;color:var(--ink-2)}
.changelog{list-style:none;margin:0;padding:0}
.changelog li{padding:16px 0;border-top:1px solid var(--line-2)}
.changelog time{font:600 13px var(--sans);color:var(--pollen-ink)}
.changelog p{margin:6px 0 0;color:var(--ink-2)}

/* ---------- Articles ---------- */
.art-grid{display:grid;grid-template-columns:repeat(auto-fill,minmax(280px,1fr));gap:32px 24px}
.art-card{text-decoration:none;display:block}
.art-card .vitrine{aspect-ratio:16/10}
.art-card h3{font-family:var(--serif);font-weight:400;font-size:26px;line-height:1.15;margin:14px 0 6px;letter-spacing:-.01em}
.art-card p{margin:0;color:var(--ink-2);font-size:15.5px}
.art-card:hover h3{text-decoration:underline;text-decoration-color:var(--pollen)}
.art-head{max-width:820px}
.art-head h1{font-size:clamp(40px,5.2vw,76px)}
.art-note{font-size:14px;color:var(--ink-3);margin:-20px 0 32px}
.art-fig{margin:0 0 clamp(36px,5vw,64px)}
.art-fig .vitrine{aspect-ratio:21/9}
.art-fig figcaption{font-size:13px;color:var(--ink-3);margin-top:10px}
.art-body{display:grid;grid-template-columns:minmax(0,8fr) minmax(0,3fr);gap:clamp(24px,5vw,80px);align-items:start}
.art-sec h2{font-family:var(--serif);font-weight:400;font-size:clamp(26px,2.6vw,34px);letter-spacing:-.015em;margin:0 0 14px}
.art-sec{margin:0 0 clamp(32px,4vw,48px)}
.art-sec p{font-size:19px;line-height:1.7;margin:0;max-width:66ch;color:var(--ink)}
.cite{font:600 11px/1 var(--sans);vertical-align:super;text-decoration:none;color:var(--pollen-ink);margin:0 3px 0 2px;white-space:nowrap}
.cite:hover{text-decoration:underline}
.cite.ev-question,.cite.ev-reported,.cite.ev-estimate,.cite.ev-observation{border:1px solid currentColor;padding:1px 4px;border-radius:999px}
.art-aside{position:sticky;top:96px}
.ev-legend{list-style:none;margin:12px 0 0;padding:0;display:grid;gap:10px}
@media (max-width:900px){.art-body{grid-template-columns:1fr}.art-aside{position:static;order:-1}.art-sec p{font-size:17.5px}}

/* ---------- languages and scripts ---------- */
:root:lang(ru){--sans:"Hanken Grotesk",ui-sans-serif,system-ui,-apple-system,"Segoe UI",Roboto,"Helvetica Neue",Arial,sans-serif}
:root:lang(zh){--serif:"Newsreader","Songti SC","STSong","Noto Serif SC","Source Han Serif SC",serif;--sans:"Hanken Grotesk","PingFang SC","Hiragino Sans GB","Microsoft YaHei","Noto Sans SC",sans-serif}
:root:lang(ja){--serif:"Newsreader","Hiragino Mincho ProN","Yu Mincho","Noto Serif JP",serif;--sans:"Hanken Grotesk","Hiragino Sans","Yu Gothic","Meiryo","Noto Sans JP",sans-serif}
:root:lang(ko){--serif:"Newsreader","AppleMyungjo","Nanum Myeongjo","Noto Serif KR",serif;--sans:"Hanken Grotesk","Apple SD Gothic Neo","Malgun Gothic","Noto Sans KR",sans-serif}
:root:lang(ar){--serif:"Noto Naskh Arabic","Geeza Pro","Arabic Typesetting","Times New Roman",serif;--sans:"Noto Sans Arabic","Geeza Pro","Segoe UI",Tahoma,sans-serif}
:root:lang(hi){--serif:"Noto Serif Devanagari","Kohinoor Devanagari","Nirmala UI",serif;--sans:"Noto Sans Devanagari","Kohinoor Devanagari","Nirmala UI",Mangal,sans-serif}
/* scripts without true italics or with joined letters: no slanting, no tracking */
:lang(zh) em,:lang(ja) em,:lang(ko) em,:lang(ar) em,:lang(hi) em{font-style:normal;color:var(--pollen-ink)}
:lang(zh) :is(h1,h2,h3,.h2,.h3,.eyebrow,.step-verb),:lang(ja) :is(h1,h2,h3,.h2,.h3,.eyebrow,.step-verb),:lang(ko) :is(h1,h2,h3,.h2,.h3,.eyebrow,.step-verb),
:lang(ar) :is(h1,h2,h3,.h2,.h3,.eyebrow,.step-verb,.btn),:lang(hi) :is(h1,h2,h3,.h2,.h3,.eyebrow,.step-verb){letter-spacing:0}
:lang(ar) :is(.eyebrow,.step-verb),:lang(hi) :is(.eyebrow,.step-verb){text-transform:none}
:lang(ar) body,:lang(hi) body{line-height:1.75}
:lang(ar) :is(h1,.h2),:lang(hi) :is(h1,.h2){line-height:1.3}
:lang(zh) body,:lang(ja) body,:lang(ko) body{line-break:strict;word-break:normal;overflow-wrap:anywhere}
:lang(ko) body{word-break:keep-all}
/* Latin names stay italic and left-to-right inside any script */
i,.latin{unicode-bidi:isolate}
[dir="rtl"] :is(i,.latin){direction:ltr}
[dir="rtl"] .btn svg,[dir="rtl"] .xlink svg,[dir="rtl"] .rail-nav svg{transform:scaleX(-1)}
[dir="rtl"] .btn:hover svg{transform:scaleX(-1) translateX(3px)}
[dir="rtl"] a.way:hover h3{transform:translateX(-10px)}
.lang-pop a:hover{background:var(--paper-2)}

/* ---------- motion layer: page transitions, image reveals, comb, loupe ---------- */
@media (prefers-reduced-motion:no-preference){
  @view-transition{navigation:auto}
  ::view-transition-old(root){animation:vt-out .28s var(--ease) both}
  ::view-transition-new(root){animation:vt-in .42s var(--ease) both}
  .hdr{view-transition-name:hdr}
}
@keyframes vt-out{to{opacity:0;transform:translateY(-8px)}}
@keyframes vt-in{from{opacity:0;transform:translateY(12px)}}
/* photographs open like a drawer in a specimen cabinet */
@supports (animation-timeline:view()){
  @media (prefers-reduced-motion:no-preference){
    :is(.sp-card,.art-card,.caste,.credit,.sp-second,.art-fig,.hive-photo,.div-top figure) .vitrine{animation:drawer linear both;animation-timeline:view();animation-range:entry 0% entry 70%}
    .comb-wrap{animation:combin linear both;animation-timeline:view();animation-range:entry 10% cover 40%}
  }
}
@keyframes drawer{from{clip-path:inset(0 0 100% 0);opacity:.4}to{clip-path:inset(0 0 0 0);opacity:1}}
@keyframes combin{from{opacity:.2;transform:scale(.96)}to{opacity:1;transform:none}}
/* comb cells fill in rings from the centre once the comb is in view */
@media (prefers-reduced-motion:no-preference){
  .comb.is-filling .cell{animation:cellin .7s var(--ease) backwards;animation-delay:calc(var(--d,0) * 1ms)}
}
@keyframes cellin{from{opacity:0;transform:scale(.4)}to{opacity:1;transform:none}}
.comb .cell{transform-box:fill-box;transform-origin:center}
/* loupe for specimen photographs (fine pointers only) */
.loupe{position:absolute;width:190px;height:190px;border-radius:50%;pointer-events:none;border:2px solid var(--pollen);
  box-shadow:0 18px 40px -12px rgba(0,0,0,.6),inset 0 0 0 1px rgba(255,255,255,.25);background-repeat:no-repeat;opacity:0;
  transform:translate(-50%,-50%) scale(.6);transition:opacity .2s,transform .25s var(--ease);z-index:3}
.vitrine.has-loupe:hover .loupe{opacity:1;transform:translate(-50%,-50%) scale(1)}
.vitrine.has-loupe{cursor:zoom-in}
.loupe-hint{position:absolute;inset-block-start:12px;inset-inline-end:12px;z-index:3;font:600 11px var(--sans);letter-spacing:.08em;text-transform:uppercase;
  color:var(--vitrine-ink);background:rgba(5,7,6,.55);padding:6px 9px;border-radius:999px;pointer-events:none;transition:opacity .3s}
.vitrine.has-loupe:hover .loupe-hint{opacity:0}
@media (prefers-reduced-motion:reduce){.loupe{transition:none}}

/* ---------- life of a honey bee (/life/) ---------- */
.life-hero{position:relative;min-height:calc(100dvh - 68px);display:grid;align-items:center;background:var(--vitrine);color:var(--vitrine-ink);overflow:hidden}
.life-hero .wrap{position:relative;z-index:1;width:100%;padding-block:clamp(56px,9vw,120px)}
.life-hero h1{font:400 clamp(46px,8.4vw,128px)/1.02 var(--serif);letter-spacing:-.025em;margin:0 0 28px;max-width:12ch;color:var(--vitrine-ink)}
.life-hero h1 em{color:var(--pollen)}
.life-hero .lede{font-size:clamp(18px,1.7vw,22px);line-height:1.5;color:var(--vitrine-ink-2);max-width:40ch;margin:0 0 18px}
.life-hero .eyebrow{color:var(--pollen)}
.life-scope{font-size:14px;color:var(--vitrine-ink-2);max-width:52ch;margin:0 0 36px;border-inline-start:2px solid var(--pollen);padding-inline-start:12px}
.life-hero-cell{position:absolute;inset-inline-end:-6vw;top:50%;width:min(62vw,820px);aspect-ratio:1;transform:translateY(-50%);color:rgba(242,183,5,.32)}
.life-hero-cell svg{width:100%;height:100%}
.life-hero-cell ellipse{transform-box:fill-box;transform-origin:50% 100%}
@media (prefers-reduced-motion:no-preference){
  .life-hero-cell path{stroke-dasharray:520;stroke-dashoffset:520;animation:hexdraw 2.4s var(--ease) .2s forwards}
  .life-hero-cell ellipse{animation:egglay 1.4s var(--ease) 1.6s both}
  .life-hero h1,.life-hero .lede,.life-hero .eyebrow,.life-scope,.life-hero .btn{animation:rise 1s var(--ease) both}
  .life-hero .lede{animation-delay:.12s}.life-scope{animation-delay:.2s}.life-hero .btn{animation-delay:.28s}
}
@keyframes hexdraw{to{stroke-dashoffset:0}}
@keyframes egglay{from{opacity:0;transform:scaleY(.2)}to{opacity:1;transform:none}}
@media (max-width:760px){.life-hero-cell{width:120vw;inset-inline-end:-50vw;opacity:.6}}

.life-story{background:var(--paper)}
.life-grid{display:grid;grid-template-columns:minmax(0,5fr) minmax(0,7fr);gap:clamp(24px,5vw,90px)}
.chaps{padding-block:12vh 30vh}
.chap{min-height:78vh;display:flex;flex-direction:column;justify-content:center;max-width:46ch;opacity:.32;transition:opacity .5s var(--ease)}
.chap.is-on{opacity:1}
.chap-n{display:flex;gap:14px;align-items:baseline;font:600 12px/1 var(--sans);letter-spacing:.12em;text-transform:uppercase;color:var(--ink-3);margin:0 0 16px}
.chap-n span{font:400 15px/1 var(--serif);letter-spacing:0;color:var(--pollen-ink)}
.chap h2{font:400 clamp(30px,3.2vw,46px)/1.08 var(--serif);letter-spacing:-.015em;margin:0 0 18px}
.chap p{font-size:18px;line-height:1.65;color:var(--ink-2);margin:0}
.life-mob{display:none}
.life-stage{position:sticky;top:84px;height:calc(100dvh - 104px);display:flex;flex-direction:column;gap:16px;padding-block:20px}
.life-vit{flex:1;border-radius:2px}
.cellsec{position:absolute;inset:0;width:100%;height:100%;padding:8% 6% 16%;transition:opacity .8s var(--ease),transform 1s var(--ease)}
.cellsec g{transition:opacity .7s var(--ease),transform .9s var(--ease);transform-box:view-box}
.cs-egg,.cs-jelly,.cs-larva,.cs-prepupa,.cs-pupa,.cs-cap,.cs-capopen,.cs-cocoon{opacity:0}
[data-stage="egg"] .cs-egg{opacity:1}
[data-stage="larva"] .cs-jelly{opacity:calc(1 - var(--p,0) * .6)}
[data-stage="larva"] .cs-larva{opacity:1;transform:translate(calc(150px * (1 - (.35 + var(--p,0) * .65))),calc(150px * (1 - (.35 + var(--p,0) * .65)))) scale(calc(.35 + var(--p,0) * .65))}
[data-stage="capping"] .cs-prepupa,[data-stage="capping"] .cs-cap,[data-stage="capping"] .cs-cocoon{opacity:1}
[data-stage="pupa"] .cs-pupa,[data-stage="pupa"] .cs-cap,[data-stage="pupa"] .cs-cocoon{opacity:1}
[data-stage="emergence"] .cs-pupa,[data-stage="emergence"] .cs-capopen{opacity:1}
[data-stage="emergence"] .cs-pupa{transform:translateX(calc(var(--p,0) * 150px))}
.cs-pupa .eye{fill:#fffdf6;transition:fill .6s}
[data-stage="pupa"] .cs-pupa .eye{fill:color-mix(in oklab,#fffdf6,#3a2410 calc(var(--p,0) * 100%))}
[data-stage="pupa"] .cs-pupa :is(.ab,.th,.hd){fill:color-mix(in oklab,#f3ecda,#8a6a3a calc(max(var(--p,0) - .55,0) * 160%))}
[data-stage="emergence"] .cs-pupa .eye{fill:#2a1a0b}
[data-stage="emergence"] .cs-pupa .ab{fill:#7a5a2a}
[data-stage="emergence"] .cs-pupa :is(.th,.hd){fill:#4a3416}
[data-stage="emergence"] .cs-pupa .stripes{stroke:#2a1a0b}
[data-stage="emergence"] .cs-pupa :is(.legs,.wingpad){stroke:#3a2812}
.life-photos .lp{position:absolute;inset:0;opacity:0;transition:opacity .9s var(--ease)}
.life-photos .lp img{transform:scale(1.06);transition:transform 6s linear}
.life-stage[data-adult] .cellsec{opacity:0;transform:scale(.92)}
.life-stage:not([data-adult]) .life-photos{visibility:hidden}
.life-photos .lp.is-on{opacity:1}
.life-photos .lp.is-on img{transform:scale(1)}
.day{position:absolute;inset-block-end:18px;inset-inline-start:22px;z-index:2;display:flex;flex-direction:column;color:var(--vitrine-ink);text-shadow:0 2px 18px rgba(0,0,0,.6)}
.day small{font:600 11px/1 var(--sans);letter-spacing:.16em;text-transform:uppercase;color:var(--pollen)}
.day b{font:400 clamp(56px,7vw,104px)/1 var(--serif);font-variant-numeric:tabular-nums;letter-spacing:-.03em}
.life-stage.no-day .day{opacity:.0}
.life-note{position:absolute;inset-block-end:16px;inset-inline-end:20px;z-index:2;margin:0;max-width:30ch;text-align:end;font-size:12px;color:var(--vitrine-ink-2)}
.tl-track{position:relative;height:44px;border-top:1px solid var(--line-2)}
.tl-seg{position:absolute;top:-4px;height:7px;inset-inline-start:calc(var(--a) * 100%);width:calc((var(--b) - var(--a)) * 100%);background:var(--paper-3);border-inline-end:2px solid var(--paper);transition:background .4s}
.tl-seg i{position:absolute;top:14px;inset-inline-start:0;font:500 11px/1.2 var(--sans);font-style:normal;color:var(--ink-3);white-space:nowrap;opacity:0;transition:opacity .3s}
.tl-seg.is-on{background:var(--pollen)}
.tl-seg.is-on i{opacity:1;color:var(--ink)}
.tl-dot{position:absolute;top:-8px;inset-inline-start:calc(var(--x,0) * 100%);width:15px;height:15px;margin-inline-start:-7px;border-radius:50%;background:var(--ink);box-shadow:0 0 0 4px var(--paper)}
.tl-ends{display:flex;justify-content:space-between;font-size:12px;color:var(--ink-3)}
@media (prefers-reduced-motion:reduce){.cellsec,.cellsec g,.life-photos .lp,.life-photos .lp img,.chap{transition:none}}
@media (max-width:900px){
  .life-grid{grid-template-columns:1fr}
  .life-stage{order:-1;top:60px;height:44svh;z-index:5;background:var(--paper);padding-block:8px 0;margin-inline:calc(var(--gut) * -1);padding-inline:var(--gut)}
  .chaps{padding-block:4vh 20vh}
  .chap{min-height:70svh;opacity:1}
  .chap p{font-size:17px}
  .tl-seg i{display:none}
  .life-note{display:none}
  .day b{font-size:52px}
}
.life-castes{background:var(--paper-2)}
.caste-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:clamp(18px,3vw,40px)}
.caste-d{margin:0}
.caste-d .vitrine{aspect-ratio:4/3}
.caste-d h3{font:400 30px/1.1 var(--serif);margin:18px 0 12px}
.cbar{height:8px;background:var(--paper-3);margin:0 0 10px;overflow:hidden}
.cbar span{display:block;height:100%;width:calc(var(--w) * 100%);background:var(--pollen);transform-origin:0 50%}
[dir="rtl"] .cbar span{transform-origin:100% 50%}
@media (prefers-reduced-motion:no-preference){.caste-d .cbar span{transform:scaleX(0);transition:transform 1.4s var(--ease)}.caste-d.is-in .cbar span{transform:none}}
.cdays{font-size:15px;color:var(--ink-2);margin:0 0 12px}
.cdays b{font:400 26px/1 var(--serif);color:var(--ink);margin-inline-end:4px}
.caste-d p{font-size:16px;color:var(--ink-2)}
@media (max-width:900px){.caste-grid{grid-template-columns:1fr}}
.life .sp-src{padding-block:clamp(48px,6vw,90px)}
.life-link{margin-top:28px;font-size:18px}
@media (max-width:900px){.cellsec{padding:6% 2% 26%}}
.tl-seg.tl-end i{inset-inline-start:auto;inset-inline-end:0}
.tl-seg[style*="--a:1.0000"]{min-width:3px}
.tl{overflow-x:clip;padding-top:8px;margin-top:-8px}
:lang(ja) :is(h1,h2,h3,.h2,.lede),:lang(zh) :is(h1,h2,h3,.h2){word-break:auto-phrase;text-wrap:balance}
:is(:lang(ja),:lang(zh),:lang(ko)) .hero h1{font-size:clamp(40px,5.6vw,84px);line-height:1.15}
:is(:lang(ja),:lang(zh),:lang(ko)) .life-hero h1{font-size:clamp(40px,6.4vw,96px);line-height:1.15}
.menu-btn,.lang>button,.btn,.chip{white-space:nowrap;overflow-wrap:normal;word-break:keep-all}